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Like Schrode said, I've used em both ways and never seemed to have any trouble. I tend to run mine so that any potential debris would get trapped in the section that can be cleaned out.If you take it apart you'll be able to see that filter element is like a cup. I've always ran mine where the fuel goes into the filter, but I don't think it really matters.
I've always run them so the crud collects on the OUTSIDE of the cup where you can see it.
Okay, I meant to say it's how I display them.Liar, we all know that you don't RUN anything.